Waddington (in his "Summary Discussion") hits the nail so square on the head that I will quote his remarks at great length: The point was made that to account for some evolutionary changes in hemoglobin, one requires about 120 amino acid substitutions...as individual events, as though it is necessary to get one of them done and spread throughout the whole population before you could start processing the next one...[and] if you add up the time for all those sequential steps, it amounts to quite a long time.

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